Nymphalis antiopa (Linnaeus, 1758)


Nymphalis antiopa: Adult [N] Nymphalis antiopa: Adult [N] Nymphalis antiopa: Adult [S] Nymphalis antiopa: Adult [S] Nymphalis antiopa: Adult (S-Germany, SW-Bavaria, 2018) Nymphalis antiopa: Adult (S-Germany, SW-Bavaria, 2018) Nymphalis antiopa: Upper side, in the spring the yellow brim has faded (Ticino) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 29. April 2022) [N] Nymphalis antiopa: Male in spring (N-Italy, Val Müstair/Val Monastero, 01. April 2017) [N] Nymphalis antiopa: Adult [S] Nymphalis antiopa: Lower side [S] Nymphalis antiopa: Adult (S-Germany, SW-Bavaria, 2018) Nymphalis antiopa: Adult (S-Germany, SW-Bavaria, 2018) Nymphalis antiopa: Female in spring (S-Germany, Isny, 18. May 2020) [N] Nymphalis antiopa: Worn female in spring (S-Germany, Isny, 18. May 2020) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 21. May 2020) [N] Nymphalis antiopa: Male in spring (S-Germany, Isny, 21. May 2020) [N] Nymphalis antiopa: Male in spring using a high willow as raised position (S-Germany, Isny, 21. May 2020) [N] Nymphalis antiopa: Male in spring using an alder tree as raised position (S-Germany, Isny, 21. May 2020) [N] Nymphalis antiopa: Eggs (N-Germany, Wolfsburg, 30. May 2020) [N] Nymphalis antiopa: Eggs (N-Germany, Wolfsburg, 30. May 2020) [N] Nymphalis antiopa: Eggs (N-Germany, Wolfsburg, 30. May 2020) [N] Nymphalis antiopa: Empty eggs with feeding traces of the L1 in surrounding leaves (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Empty eggs (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Empty eggs (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [N] Nymphalis antiopa: Larvae L1 in the end of this first instar (S-Germany, Isny, 17. June 2023) [M] Nymphalis antiopa: Larvae L1 in the end of this first instar, one larva already in L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L1 in the end of this first instar, some larvae already in L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L1 in the end of this first instar, some larvae already in L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2, one still in L1 (S-Germany, Isny, 17. Juni 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, 17. June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L2 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L2 in the end of this instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2023) Nymphalis antiopa: Larvae L3 (S-Germany, Isny, June 2018) [N] Nymphalis antiopa: Half-grown larvae L3 (S-Germany, Isny, Adelegg, June 2018) [N] Nymphalis antiopa: Half-grown larva L3 (S-Germany, Isny, Adelegg, June 2018) [N] Nymphalis antiopa: Half-grown larva L3 (S-Germany, Isny, Adelegg, June 2018) [N] Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L4, penultimate instar (S-Germany, Isny, June 2023) Nymphalis antiopa: Larva L5 (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva [S] Nymphalis antiopa: Larvae [S] Nymphalis antiopa: Larvae [S] Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) [S] Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larvae [S] Nymphalis antiopa: Larvae [S] Nymphalis antiopa: Larva Nymphalis antiopa: Larva [S] Nymphalis antiopa: Larva [S] Nymphalis antiopa: Larvae [S] Nymphalis antiopa: Larva [S] Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Larva (S-Germany, Isny, Adelegg, June 2018) Nymphalis antiopa: Pupa (S-Germany, Isny, Adelegg, 2018) Nymphalis antiopa: Pupa (S-Germany, Isny, Adelegg, 2018) Nymphalis antiopa: Pupa [S] Nymphalis antiopa: Pupa [S] Nymphalis antiopa: Pupa [S] Nymphalis antiopa: Pupa [S] Nymphalis antiopa: Habitat in the Black forest in southwest Germany in June 2009: clearcutting and storm successions with Salix and Betula in large, mountainous forests [N] Nymphalis antiopa: Habitat in the Adelegg (a prealpine lower massif in Southern Germany) in a sunny stream valley with Salix between mountainous forests (June 2010) [N] Nymphalis antiopa: Larvae L1 in their nest and feeding traces (S-Germany, Isny, 17. Juni 2023) [N] Nymphalis antiopa: Larvae L1 in their nest and feeding traces (S-Germany, Isny, 17. Juni 2023) [N] Nymphalis antiopa: Larvae on Salix alba at an atypical place in 1000m in the northern foothills of the Alps (S-Germany, Adelegg, June 2018) [N] Nymphalis antiopa: Larvae on Salix alba at an atypical place in 1000m in the northern foothills of the Alps (S-Germany, Adelegg, June 2018) [N] Nymphalis antiopa: Larvae on Salix appendiculata (S-Germany, Isny, Adelegg, June 2018) [N] Nymphalis antiopa: Larval habitat with Salix appendiculata (S-Germany, Isny, Adelegg, June 2018) [N] Nymphalis antiopa: Habitat (S-Germany, Isny, 18. May 2020) [N] Nymphalis antiopa: Habitat (S-Germany, Isny, 18. May 2020) [N]

Host plants:
The larvae feed on Salicaceae (especially willows like Salix caprea) and Betulaceae (birches like Betula pendula or Betula pubescens), rarely probably also on other woods.

Habitat:
Nymphalis antiopa inhabits large forests, particularly in cool, moist, mountainous areas. It is rarely found in parks, especially in years with higher abundance.

Life cycle:
Nymphalis antiopa occurs in a single generation per year. Overwintered adults fly from April to June, sometimes even until early or mid-July. Oviposition takes place much later than in Nymphalis polychloros (according to year and altitude mostly between mid-May and early June, on twigs mostly in some hight). The egg phase seems to last longer than in N. polychloros, too, resulting in larvae about one month later than in polychloros. The caterpillars can be found from June to August, often in larger communities. The resulting butterflies appear in August/September.

Endangerment: endangered

Endangerment factors:
Nymphalis antiopa is endangered through monotonization and darkening of the forests, removal of economically worthless soft woods, dense coniferous afforestation and forest fragmentation. Nymphalis antiopa itself has also pronounced cycles of abundance (possibly influenced by parasitoids). Nymphalis antiopa was quite common on the Adelegg (a prealpine low mountain range in southern Germany) between 1990-1996. After this period Nymphalis antiopa was very rare there again. First new own observations succeeded not before 20010 and 2011 (from 2020-2023 common there again). Elsewhere, such as in the Upper Bavarian Alps Nymphalis antiopa was observed regularly in the years 2004-2007 (Mario Peluso, Ulrich Rau).

Remarks:
The distribution is Holarctic. Nymphalis antiopa occurs from northern Spain through Europe and temperate Asia to North America.



Nymphalis c-album | Nymphalis egea | Nymphalis ichnusa | Nymphalis io | Nymphalis polychloros | Nymphalis urticae | Nymphalis xanthomelas