Tarucus balkanicus (Freyer, 1844)


Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Male (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Imago (N-Greece, Askio mountains near Siatista, 02. June 2021) [N] Tarucus balkanicus: Female (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Female (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Female (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Female (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Female (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Adult lower side (Askio mountains. N-Greece, Kozani, May 2008) [M] Tarucus balkanicus: Adult (Askion, July 2011) [N] Tarucus balkanicus: Imago (Greece, Siatista near Kozani, late June 2013) [N] Tarucus balkanicus: Upper side (Kozani, N-Greece, May 2008) [N] Tarucus balkanicus: Upper side (Kozani, N-Greece, May 2008) [N] Tarucus balkanicus: Lower side (Kozani, May 2008) [N] Tarucus balkanicus: Ovum at base of a thorn (July 2011, Askion) [N] Tarucus balkanicus: Ovum at base of a thorn (July 2011, Askion) [N] Tarucus balkanicus: The eggs are sometimes also deposited on the leaves (Askio mountains near Siatista, early August 2012) [N] Tarucus balkanicus: Egg (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: Egg (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: Egg (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: Egg, laid on two already hatched egg shells (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: Empty egg shell and old feeding scars of a larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: L1 (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L1 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L2 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 and L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L3 and two L4 larvae (e.o., N-Greece, Askio mountains near Siatista, eggs found in mid-July 2025) [S] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [S] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [M] Tarucus balkanicus: L4 larva (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: Larva in L4 (Askion, July 2011) [M] Tarucus balkanicus: Larva (Askio mountains, Greece, early August 2012) [N] Tarucus balkanicus: Larva in L4 (Askion, July 2011) [M] Tarucus balkanicus: Larva in L4 (Askion, July 2011) [M] Tarucus balkanicus: Präpupa [S] Tarucus balkanicus: Prepupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa (e.l., N-Greece, Askio mountains near Siatista, larvae found in mid-July 2025) [S] Tarucus balkanicus: Pupa [S] Tarucus balkanicus: Feeding pattern at a leaf (Askion, July 2011) [N] Tarucus balkanicus: Infested twig directly at a rock (Askion, July 2011) [N] Tarucus balkanicus: Habitat in the north Greek Askio mountains near Siatista, July 2011. I observed also Pieris ergane and Chilades trochylus. [N] Tarucus balkanicus: Habitat in Northern Greece (Kozani, Siatista; May 2008) [N] Tarucus balkanicus: Habitat (N-Greece, Askio mountains near Siatista, mid-July 2025) [N] Tarucus balkanicus: Habitat (N-Greece, Askio mountains near Siatista, mid-July 2025) [N]

Host plants:
The caterpillars live on Paliurus spina-christi (Rhamnaceae). This plant is favoured by grazing.

Habitat:
Tarucus balkanicus inhabits hot and dry sheep pastures, rocky slopes and similar grassland areas where the host plant (a bramble) is frequent.

Life cycle:
Tarucus balkanicus has several generations per year. I met adults of the first generation in Northern Greece in early May at the foot of slopes. In late July 2011 and in mid-July 2025 I found caterpillars and eggs in the Askio Mountains (northern Greece) quite numerous at an extremely xerothermic slope in heat accumulation over stones, rocks and niches. Eggs are laid at the base of spines, rarely also on leaves. The larva has a characteristic feeding pattern with one to four mines strips in the longitudinal direction on the leaf. Larvae are often accompanied by ants. Pupation occurs mostly on stems or in the litter (e.g. on old dry leaves).

Endangerment factors:
Tarucus balkanicus is threatened by habitat changes and overbuilding (e.g. east of Mount Olympus). Looking ahead, also a threat may arise from land use through solar farms and wind parks, which may be constructed in ecologically valuable areas (Askion etc.). A major threat is finally the abandonment of grazing with sheep and goats.

Remarks:
Tarucus balkanicus flies locally in Africa and is found from the Balkans (from Croatia) across Turkey and the Arabian Peninsula to Central Asia.



Tarucus theophrastus