Tarucus theophrastus (Fabricius, 1793)


Tarucus theophrastus: Female (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Female (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Female on Zygophyllum, an important nectar source (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Female on Zygophyllum, an important nectar source (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Female (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Egg on lower side of a leaf. Near the base of the leaf another already hatched egg is visible on the stem (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Empty egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Empty egg (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Half-grown larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Half-grown larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Half-grown larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [S] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva with ants (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [M] Tarucus theophrastus: Larva (S-Spain, Andalusia, Cabo de Gata, late September 2017) [S] Tarucus theophrastus: Pupa (e.l. rearing, S-Spain, Andalusia, Cabo de Gata, late September 2017) [S] Tarucus theophrastus: Pupa (e.l. rearing, S-Spain, Andalusia, Cabo de Gata, late September 2017) [S] Tarucus theophrastus: Pupa (e.l. rearing, S-Spain, Andalusia, Cabo de Gata, late September 2017) [S] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Larval habitat with Zizyphus lotus (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N] Tarucus theophrastus: Habitat with Zizyphus lotus. One of the two major threats (the overbuilding) is clearly visible. Together with greenhouse agriculture it has made large parts of the Andalusian coast an ecological desert (S-Spain, Andalusia, Cabo de Gata, late September 2017) [N]

Host plants:
The larvae usually live on Zizyphus lotus (Rhamnaceae), so in Spain and Morocco, but in other parts of the distribution also on related plants such as Paliurus.

Habitat:
In Spain, Tarucus theophrastus inhabits sandy or stony, coastal places such as dunes.

Life cycle:
In Spain, the butterflies occur from April to winter. Oviposition usually takes place on the lower side of the leaves, partly also on the stem near a leaf or at the base of thornes. The larvae cause characteristic feeding scars (longituninal stripes). They are usually accompanied by ants and often rest on the lower side of leaves near the ground. The pupa is located on the ground or larger stems.

Endangerment factors:
Tarucus theophrastus is heavily endangered in Europe. The typical endangerment factors overbuilding (tourism etc.) and intense EU-agriculture (e.g. greenhouses) have left only very few sites.

Remarks:
Tarucus theophrastus occurs in Africa and the Arabian Peninsula. In Europe it is restricted to a few small, isolated populations at the Spanish Mediterranean coast (especially Almeria, even more sporadically also Murcia and Granada).



Tarucus balkanicus