Cupido minimus (Fuessly, 1775)


Cupido minimus: Males (S-Germany, Allgäu Alps, 1600m, 22. June 2014) [N] Cupido minimus: Adult (Switzerland, Simplon, early July 2019) [N] Cupido minimus: Female (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Male (S-Germany, Allgäu, Oberjoch, 20. May 2020) [N] Cupido minimus: Male and copulation [N] Cupido minimus: Male tries to disturb other mating [N] Cupido minimus: Adult [N] Cupido minimus: Female [N] Cupido minimus: Male (Askion, Northern Greece, May 2010) [N] Cupido minimus: Upper side [N] Cupido minimus: Lower side [S] Cupido minimus: Mud-puddling males, together with tages, semiargus and orbitulus  (S-Germany, Allgäu Alps, 1600m, 22. June 2014) [N] Cupido minimus: Copulation (S-Germany, Allgäu Alps, Hinterstein, 1600m, 03. June 2020) [N] Cupido minimus: Mating [N] Cupido minimus: Oviposition (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Oviposition (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Oviposition (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Oviposition (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Egg on Anthyllis (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Egg on Anthyllis (S-Germany, Gerstetten, 15. June 2022) [N] Cupido minimus: Egg on Anthyllis (S-Germany, Gerstetten, 15. June 2022) [N] Cupido minimus: Egg (S-Germany, Allgäu, Oberjoch, 20. May 2020) [N] Cupido minimus: Egg (S-Germany, Allgäu, Oberjoch, 20. May 2020) [N] Cupido minimus: Egg (S-Germany, Allgäu, Oberjoch, 20. May 2020) [N] Cupido minimus: Egg on Astragalus monspessulanus (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Egg on Astragalus monspessulanus (Switzerland, Valais, Täschalpe, mid-June 2022) [N] Cupido minimus: Ovae [N] Cupido minimus: Ovum [N] Cupido minimus: Ovum, detail [N] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [M] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [M] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [M] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L1-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L2-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L2-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L2-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L2-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L3-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L4-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L4-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L4-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L4-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: L4-larva (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva in the ultimate instar (S-Germany, Gerstetten, mid-June 2022) [S] Cupido minimus: Larva [N] Cupido minimus: Larva (Kanisfluh, Bregenzer Wald, July 2010) [N] Cupido minimus: Die larvae are well camouflaged while consuming the seeds in the calyx (Kanisfluh) [S] Cupido minimus: Larva that has changed to Lotus due to lack of Anthyllis (France, southern Alps, Col de Champs, early August 2021) [M] Cupido minimus: Fully-grown larva on its way into a shelter for hibernation (S-Germany, Gerstetten, late June 2022) [S] Cupido minimus: Fully-grown larva in dormancy (S-Germany, Gerstetten, late June 2022) [S] Cupido minimus: Larva after the hibernation (found in a grass tussock, Valais 2008) [M] Cupido minimus: Larva after the hibernation (found in a grass tussock, Valais 2008) [M] Cupido minimus: Pupa [S] Cupido minimus: Pupa [S] Cupido minimus: Pupa lateral [S] Cupido minimus: Pupa ventral [S] Cupido minimus: Habitat (S-Germany, Heidenheim/Brenz, 11. June 2006) [N] Cupido minimus: Habitat (S-Germany, Heidenheim/Brenz, 11. June 2006) [N] Cupido minimus: Habitat (S-Germany, Allgäu Alps, Hinterstein, 1800m, 18 August 2012) [N]

Host plants:
Oviposition mostly occurs on Anthyllis vulneraria, rarely also Astragalus cicer (e.g. Germany, eastern Swabian Alb) and Colutea arborescens (e.g. Germany, Kaiserstuhl, according to literature). In the Alps, I observed the oviposition also on Oxytropis campestris and in June 2022 on Astragalus monspessulanus. The species also uses other Astragalus and Oxytropis species.

Habitat:
Cupido minimus inhabits nutrient-poor grasslands and extensively grazed pastures of all kinds with stocks of the host plant.

Life cycle:
Cupido minimus flies mostly in a single, more rarely and partial also two generations per year. In the lowlands, the butterflies fly mostly from mid/late May to mid-July, at high altitudes (above 2500m asl) from June to August. In 2007, I already observed first adults as early as 30th April at 1800m asl on a south facing slope in the Montafon (west Austria)! Eggs are usually deposited on calyxes. The caterpillar lives at first within the calyx and later also outside in the fruit stand. It feeds especially on the ovary and developing seeds. The fully-grown caterpillar overwinters mostly in dense grass tussocks, where also pupation occurs in spring. In Valais, I observed several hibernated larvae in Festuca-tufts in about 1500 meters above sea level in April.

Endangerment: regionally endangered or decreasing

Endangerment factors:
Where nutrient-poor grasslands are still more common, Cupido minimus is endangered only slightly. Otherwise, the butterfly is often already extinct or at least very local. In the Alps, the butterfly is the most common Lycaenidae.

Remarks:
Cupido minimus is widespread in Europe and spreads in Asia to Mongolia. The populations in southern Spain are partly considered as a separate species.



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