Pieris balcana Lorkovic, [1969]


Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Male of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female feeding on Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female feeding on Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female feeding on Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Female of the second generation (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, mid-June 2023) [S] Pieris balcana: Egg on the lower leaf side of Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [M] Pieris balcana: Egg on the lower leaf side of Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [S] Pieris balcana: Egg on the lower leaf side of Calepina irregularis (N-Greece, Pontokomi near Kozani, early April 2022) [S] Pieris balcana: Egg (N-Greece, Pontokomi near Kozani, early April 2022) [S] Pieris balcana: Egg (N-Greece, Pontokomi near Kozani, early April 2022) [S] Pieris balcana: Egg (N-Greece, Pontokomi near Kozani, early May 2023, leg. Gregor Markl) [S] Pieris balcana: Egg (N-Greece, Pontokomi near Kozani, early May 2023, leg. Gregor Markl) [S] Pieris balcana: Larva L1 next to the mostly consumed egg shell and a ready-for-hatching batch of eggs of P. brassicae on the same leaf (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023, leg. Gregor Markl) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L1 on the top, on the bottom L2 in the end of this instar (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 1 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 2 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 3 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 4 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 6 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 7 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 8 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 9 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L4 moulting into L5, pic 10 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Larva L5 (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Prepupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Prepupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Prepupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Prepupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Pupa prior to eclosion (e.o. rearing, N-Greece, Pontokomi near Kozani, oviposition in early May 2023) [S] Pieris balcana: Habitat. The host plant Calepina irregularis grew on somewhat slightly more humid spots like here on the margin of the quarry (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Habitat. The host plant Calepina irregularis grew on somewhat slightly more humid spots like here on the margin of the quarry (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: The females were only interested in Calepina irregularis. They also used this plant (among others) for nectaring (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: The females were only interested in Calepina irregularis. They also used this plant (among others) for nectaring (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Habitat (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Habitat (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Habitat (N-Greece, Pontokomi near Kozani, early April 2022) [N] Pieris balcana: Habitat (N-Greece, Pontokomi near Kozani, early April 2022) [N]

Host plants:
The larvae feed on various Brassicaceae, most often in semi-shady or shady position. I recorded oviposition in northern Greece on Calepina irregularis.

Habitat:
Pieris balcana occurs especially in edge habitats like woodland and grove margins, at the edges of Rubus stands, in abandoned quarries, in river valleys, in cultural land that is interspersed with bushes and in similar places. Contrary to P. napi it is also found in hot and dry environments provided that there are disturbed places like quarries or shrub.

Life cycle:
The adults occur in several generations between March and October. In dry regions they may be rarer in summer (drought) than in spring. In early April 2022 the adults often have spent the night within nearby Rubus thickets (GR-Kozani). Oviposition usually occurs on lower sides of leaves. The larvae develop very fast like in P. napi.

Remarks:
Pieris balcana occurs in the Balkan peninsula (from Croatia and S-Romania to especially Greece). In Greece, P. napi seems to be rare or is restricted to the very North and Northeast (has to be investigated). Pieris balcana, however, is regionally quite common. In April 2023 all checked specimens prooved balcana. It shows - in contrast to P. napi - an in average not sharply confined dark squamation of the veins (hind wing undersides). This is especially visible in the spring generation, but not in all cases very distinctly.
The distribution to the east ist poorly known. It at least occurs in S-Ukraine and S-Russia.



Pieris brassicae | Pieris bryoniae | Pieris cheiranthi | Pieris ergane | Pieris krueperi | Pieris mannii | Pieris napi | Pieris rapae