Leptidea morsei (Fenton, 1882)


Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Female (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Male (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Male, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Female, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Female, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Female, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Female, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Female, second generation (Romania, near Cluij-Napoca, oviposition in the first half of May 2021, mid-June 2021) [S] Leptidea morsei: Eggs (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Eggs (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Egg (Romania, near Cluij-Napoca, first half of May 2021) [M] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the first instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the second instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the third respectively penultimate instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Larva in the final instar (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Prepupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Pupa few days prior to eclosion (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Male pupa prior to eclosion (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Empty pupal skin (Romania, near Cluij-Napoca, oviposition in the first half of May 2021) [S] Leptidea morsei: Egg in usual position (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Plant with egg (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Plant with egg (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Plant with egg (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Plant with egg (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Oviposition habitat (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Oviposition habitat (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Oviposition habitat (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Oviposition habitat (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Oviposition habitat (Romania, near Cluij-Napoca, first half of May 2021) [N] Leptidea morsei: Habitat, oak woodland (Romania, near Cluij-Napoca, first half of May 2021) [N]

Host plants:
The larvae usually feed on Lathyrus niger, locally also on closely related other Lathyrus species.

Habitat:
Leptidea morsei is a typical inhabitant of open, light woodlands, especially warm, mesotrophic and mesophilous oak woodlands, where the host plant occurs in the understory. Oaks are light forest species which let more light through ther canopy than e.g. beech or marple. Leptidea morsei is also known from riparian woodland.

Life cycle:
This largest European Leptidea species usually occurs in very low densities and thus needs large habitats. The butterfly appears most often in two generations: the first from late April (sometimes already from mid-April) until mid-June and the second one between late June and August. In cool regions there may be a single generation in June/July only. Males patrol along forest tracks and other woodland edges. Oviposition usually takes place singly on the underside of a leaf of not too small plants. The larva develops on a single plant and does not change it. The decvelopment is fast with three larval moults only. Larvae on living plants which resulted in non-hibernation pupae pupated on the host plant without migration phase.

Endangerment factors:
Leptidea morsei is heavily endangered in eastern central Europe (e.g. E-Austria) due to abandonment of traditional forest management (e.g. mixed coppice, fuel wood), afforestation with dark forest species such as beech or maple and eutrophication. Leptidea morsei is listed in the annexes of the Flora-Fauna-Habitat directive of the European Union, but this would only be helpful if the resulting area management plans were proper translated in actual an sustainable field measures.

Remarks:
Leptidea morsei occurs from eastern central Europe (E-Austria, Czech Republic, Slovenia) eastward across E-Europe and temperate Asia to N-Japan. In Europe, it is principally a lowland species which does usually not occur above 1000m. In Asia, however, it reaches locally 2000m.



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